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PWRC Software Archive

The following programs were developed at PWRC and elsewhere for the analysis of animal populations for wildlife biologists/managers.

BAND2Computes the number of animals that must be banded in order to achieve a specified level of precision for survival rates estimated using band recovery models.
BROWNIEComputes estimates of survival and recovery rates for 2-age class band-recovery data.
CAPQUOTAComputes expected coefficients of variation of survival and capture probability for use in the design of capture-recapture-resighting experiments.
CAPTUREComputes estimates of capture probability and population size for "closed" population capture-recapture data.
CENTROIDTests the null hypothesis that two samples of recoveries belong to the same bivariate distribution.
COMDYNComputes estimates of species richness using large scale data.
CONTRASTCompares estimates of survival (or any normally distributed parameter estimates).
CountRemovalA removal model for estimating detection probabilities from point count surveys.
DOSECOMPAnalyzes dose responses for two populations, using the PROBIT model and the abbreviated protocol as described by Link et al(1996).
DOBSERVA Double-observer approach for estimating detection probability and abundance from avian point counts as described in Nichols et. al. (2000).
ESTIMATEComputes estimates of survival and recovery rates for 1-age class band-recovery data.
fatalityCMR Estimates fatalities using raw counts of wildlife fatalities, trial experiments for carcass detection probability and persistence time.
GENCAPH1Generates capture-history data for given s,theta,p,c.
JOLLYComputes estimates of survival and capture probability for 1-age class open population capture-recapture models.
JOLLYAGEComputes estimates of survival and capture probability for 2-age class open population capture-recapture models.
LOLASURVIVComputes estimates of survival and capture probability for open capture-recapture models when sex is not allways known.
MAYFIELDComputes estimates of daily survival rate for nest visitation data.
MSSURVIVComputes estimates of survival-movement and capture probability for open capture-recapture models.
MSSRVRD(BETA) replacement for MSSURVIV, RDSURV - Combines multi-strata, robust-design and trap-deaths.
MSSRVRCVComputes survival-movement, capture and recovery probabilities from multi-strata capture-recapture-recovery data.
MSSRVMISComputes estimates of survival and capture probability adjusted for missclassification.
MULTComputes estimates of survival and recovery rates for band-recovery data with additional models for estimating reporting rate.
nestAbund Estimates abundance of nests using a capture-recapture approach which deals with imperfect detection probability as well as the impact of environmental and individual covariates on nest fate.
ORDSURVIVComputes estimates of survival and temporary emigration under the "Open Robust Design".
POWERComputes the power of between-model likelihood ratio tests.
PRESENCE Estimates patch occupancy rates and related parameters.
RDSURVIVComputes estimates of survival and capture probability using capture-recapture data with "robust-design" models and recoveries.
RELEASEComputes estimates of survival and capture probability for capture-recapture experiments on open animal populations.
SCATMANTests the hypothesis of prey-selectivity based on random samples of predator scats.
SPACECAP Estimates Animal Abundance and Density using Spatially-Explicit Capture-Recapture.
SPECRICHComputes "species richness" or total number of species from emperical species abundance distribution data.
SPECRICH2Estimates the total number of species from species presence-absence data on multiple sample sites or occasions using model M(h) from program CAPTURE.
SURVIVComputes estimates of survival (or any model parameter) with multinomially distributed data.
TMSURVIVComputes estimates of survival and capture probability and the proportion of transients in open population capture-recapture data.

Other www sites with wildlife-related software:


Although this program has been used by the U.S. Geological Survey (USGS), no warranty, expressed or implied, is made by the USGS or the U.S. Government as to the accuracy and functioning of the program and related program material nor shall the fact of distribution constitute any such warranty, and no responsibility is assumed by the USGS in connection therewith.


One of the problems with obtaining software from the Web is that hard copy documentation is not available. The following site provides information on how to cite electronic documents:

Here are sample citations for two of the programs:

Program SURVIV requires a FORTRAN compiler to generate the user-defined estimation routine. If you do not have one, click here to download the FREE Gnu FORTRAN compiler for WINDOWS.

A good, free ascii text editor (Notepad++) can be downloaded here.

This list is maintained by:

Jim Hines (jhines@usgs.gov)
Patuxent Wildlife Research Center
12100 Beech Forest Rd
Laurel, Md 20708-4030

Last updated: 31-May-2017